Understanding “What in pollination?” meme

I came across many images on Twitter like this

update 1
Original link.

Does the hat resemble nation? If yes then what’s the reason?
I came to know that everything that ends with nation or sounds like nation they use this sticker and laugh at it.
But I don’t get the joke.

Answer

The joke is that this is a parody of another popluar meme.
From knowyourmeme.com, What in Tarnation,
enter image description here

“When you find a city slicker on FarmersOnly.com / wot n ternation”

About
“What in Tarnation?” is a rhetorical question meaning “what in damnation?”, which is often associated with Americans living in the Southern United States expressing incredulous bewilderment.

The meme above spawned many parody memes. Here are some of the memes.
enter image description here

enter image description here

Notice that the parodies have the hat in common, follow the formula what in X where X rhymes with tarnation, and the image is somehow related to X.

In the OP, notice that the image is of a bee, bees pollinate, and the act of pollinating is pollination.

Attribution
Source : Link , Question Author : Ashutosh Soni , Answer Author : Em.

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